Francisella tularensis holarctica (ex Olsufjev et al. 1959) Olsufjev and Meshcheryakova 1983 Olsufjev N.G. Meshcheryakova I.S. Subspecific taxonomy of Francisella tularensis McCoy and Chapin 1912 1983 33 4 872 874 International Journal of Systematic Bacteriology Int J Syst Bacteriol 0020-7713 10.1099/00207713-33-4-872 Svensson K. Larsson P. Johansson D. Byström M. Forsman M. Johansson A. Evolution of subspecies of Francisella tularensis. 2005 187 11 3903 8 Journal of Bacteriology J Bacteriol 0021-9193 1098-5530 10.1128/JB.187.11.3903-3908.2005 15901721 Verified that there is no sequence data available (May 1, 2018). Assigned a proxy type. Strain GIEM 503 is the attenuated live vaccine strain. Isolated from a tick in the Moscow area of Russia in 1949.