Treponema pallidum endemicum Smibert 1984 Universal Root Woese et al. 1990 Bacteria (None 2015) Woese et al. 1990 Spirochaetota corrig. Garrity and Holt 2021 Spirochaetes (sic) Garrity and Holt 2021 non Spirochaetes Cavalier-Smith 2002 Spirochaetia (Cavalier-Smith 2002) Paster 2020 Spirochaetales Buchanan 1917 (Approved Lists 1980) emend. Hördt et al. 2020 Treponemataceae Robinson 1948 (Approved Lists 1980) emend. Hördt et al. 2020 Treponema Schaudinn 1905 (Approved Lists 1980) emend. Abt et al. 2013 Treponema pallidum (Schaudinn and Hoffmann 1905) Schaudinn 1905 (Approved Lists 1980) emend. Smibert 1984 Treponema pallidum endemicum Smibert 1984 Treponema pallidum (Schaudinn and Hoffmann 1905) Schaudinn 1905 (Approved Lists 1980) emend. Smibert 1984 Treponema pallidum pallidum (Schaudinn and Hoffmann 1905) Smibert 1984 Treponema pallidum pertenue (Castellani 1905) Smibert 1984 Treponema pallidum carateum (Smibert 1984) None 2015 bejel Treponema pallidum endemicum Smibert 1984 bejel Treponema pallidum endemicum Smibert 1984 Staudová B. Strouhal M. Zobaníková M. Cejková D. Fulton L.L. Chen L. Giacani L. Centurion-Lara A. Bruisten S.M. Sodergren E. Weinstock G.M. Smajs D. Whole genome sequence of the Treponema pallidum subsp. endemicum strain Bosnia A: the genome is related to yaws treponemes but contains few loci similar to syphilis treponemes 2014 8 11 e3261 PLoS Neglected Tropical Diseases PLoS Negl Trop Dis 1935-2727 1935-2735 10.1371/journal.pntd.0003261 25375929 Cejková D. Zobaníková M. Pospísilová P. Strouhal M. Mikalová L. Weinstock G.M. Smajs D. Structure of rrn operons in pathogenic non-cultivable treponemes: sequence but not genomic position of intergenic spacers correlates with classification of Treponema pallidum and Treponema paraluiscuniculi strains 2013 62 2 196 207 Journal of Medical Microbiology J Med Microbiol 0022-2615 1473-5644 10.1099/jmm.0.050658-0 23082031 bejel Treponema pallidum endemicum Smibert 1984 Mikalová L. Strouhal M. Oppelt J. Grange P.A. Janier M. Benhaddou N. Dupin N. Šmajs D. Human Treponema pallidum 11q/j isolate belongs to subsp. endemicum but contains two loci with a sequence in TP0548 and TP0488 similar to subsp. pertenue and subsp. pallidum, respectively 2017 11 3 e0005434 PLoS Neglected Tropical Diseases PLoS Negl Trop Dis 1935-2727 1935-2735 10.1371/journal.pntd.0005434 28263990 Cejková D. Zobaníková M. Pospísilová P. Strouhal M. Mikalová L. Weinstock G.M. Smajs D. Structure of rrn operons in pathogenic non-cultivable treponemes: sequence but not genomic position of intergenic spacers correlates with classification of Treponema pallidum and Treponema paraluiscuniculi strains 2013 62 2 196 207 Journal of Medical Microbiology J Med Microbiol 0022-2615 1473-5644 10.1099/jmm.0.050658-0 23082031 Smibert R.M. Holt J.G. Genus III. Treponema Schaudinn 1905, 1728(AL) 1984 1 1 49 57 Bergey's Manual of Systematic Bacteriology The Williams and Wilkins Co. Baltimore