Hydrogenosomobacter Takeshita et al. 2019 Hydrogenosomobacter endosymbioticus Takeshita et al. 2019 Universal Root Woese et al. 1990 Bacteria (None 2015) Woese et al. 1990 Proteobacteria (sic) Garrity et al. 2021 Pseudomonadota corrig. Garrity et al. 2021 Alphaproteobacteriota (sic) Garrity et al. 2021 Alphaproteobacteria Garrity et al. 2006 Holosporales Szokoli et al. 2020 Holosporaceae Görtz and Schmidt 2006 emend. Szokoli et al. 2016 Hydrogenosomobacter Takeshita et al. 2019 Holosporaceae Görtz and Schmidt 2006 emend. Szokoli et al. 2016 Holospora (ex Hafkine 1890) Gromov and Ossipov 1981 Goertzia corrig. Boscaro et al. 2013 Gortzia (sic) Boscaro et al. 2013 Paraholospora Eschbach et al. 2009 Hepatobacter Nunan et al. 2013 Bealeia Szokoli et al. 2016 Hafkinia Fokin et al. 2019 Cytomitobacter Tashyreva et al. 2018 Nesciobacter (sic) George et al. 2020 Ignotibacter corrig. George et al. 2020 Fujishimia Boscaro et al. 2019 Hydrogenosomobacter endosymbioticus Takeshita et al. 2019 Takeshita K. Yamada T. Kawahara Y. Narihiro T. Ito M. Kamagata Y. Shinzato N. Tripartite Symbiosis of an Anaerobic Scuticociliate with Two Hydrogenosome-Associated Endosymbionts, a Holospora-Related Alphaproteobacterium and a Methanogenic Archaeon. 2019 85 24 Applied and Environmental Microbiology Appl Environ Microbiol 0099-2240 1098-5336 10.1128/AEM.00854-19 31585988